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A protein designated JASSY transfers 12oxoPDA from the chloroplast outer membrane into the cytosol, before this intermediate is transferred by an ATP-binding cassette transfer protein (designated 'CTS1' or 'PAX1') from the cytosol to the peroxisomes, where a flavin-dependent 12oxo-phytodienoate reductase (OPR3) reduces the double bond in position 10, i.e., in the cyclopentenone ring, to yield 3-oxo-2-(pent-2'-enyl)-cyclopentane-1-octanoic acid